This list comprises research publications and presentations by Badlands Dinosaur Museum staff and associated researchers, and also research by other scholars on specimens held in our collections.

Curator: Dr. Denver Fowler


Research Publications

Warshaw EA & Fowler DW (2022) A transitional species of Daspletosaurus Russell, 1970 from the Judith River Formation of eastern Montana, PeerJ. http://dx.doi.org/10.7717/peerj.14461 (Publication scheduled online 7am GMT Nov 25th 2022, free access). (Summary / Press Links / Images)

Wilson JP, & Fowler DW (2020) The easternmost occurrence of Saurornitholestes from the Judith River Formation, Montana, indicates broad biogeographic distribution of Saurornitholestes in the Western Interior of North America. Historical Biology. https://doi.org/10.1080/08912963.2020.1862828 (published online Dec. 23rd 2020) (Summary / Press Links / Images)

Fowler DW (2020) The Hell Creek Formation, Montana: A Stratigraphic Review and Revision Based on a Sequence Stratigraphic Approach. Geosciences 2020, 10(11), 435. https://doi.org/10.3390/geosciences10110435 (Free Access PDF)

Fowler DW, Wilson JP, Freedman Fowler EA, Noto CR, Anduza DA, & Horner JR (2020) Trierarchuncus prairiensis gen. et sp. nov., the last alvarezsaurid: Hell Creek Formation (uppermost Maastrichtian), Montana". Cretaceous Research 116, December 2020, 104560. doi:10.1016/j.cretres.2020.104560 (Published online July 10th 2020) (Summary / Press Links / Images)

Fowler DW & Freedman Fowler EA (2020) Transitional evolutionary forms in chasmosaurine ceratopsid dinosaurs : evidence from the Campanian of New Mexico, PeerJ 8:e9251. DOI 10.7717/peerj.9251 (Free Access PDF) (Published online June 5th 2020) (Summary / Press Links / Images)

Canoy Illies MM & Fowler DW (2020) Triceratops with a kink: Co-ossification of five distal caudal vertebraefrom the Hell Creek Formation of North Dakota. Cretaceous Research (108): https://doi.org/10.1016/j.cretres.2019.104355 (Summary / Press Links / Images)

Fowler DW (2017) Revised geochronology, correlation, and dinosaur stratigraphic ranges of the Santonian-Maastrichtian (Late Cretaceous) formations of the Western Interior of North America. PLoS ONE 12(11): e0188426. https://doi.org/10.1371/journal.pone.0188426 (Free Access PDF)

Woodruff DC, Fowler DW, & Horner JR (2017) A new multi-faceted framework for deciphering diplodocid ontogeny. Palaeontologica Electronica 20.3.43A: 1-53. https://doi.org/10.26879/674 (Free Access PDF)

Wilson JP & Fowler DW (2017) First confirmed identification of juvenile Triceratops epiparietals. Cretaceous Research 70, p. 71-76. https://doi.org/10.1016/j.cretres.2016.10.005

Hall LE, Hall AE, & Fowler DW (2016) Using the trackway fossil record to test the substrate grip hypothesis for sauropod pedal unguals. In: Richter A, and Manning PL (eds.), Dinosaur Tracks - next steps, Indiana University Press.

Wings O, Tutken T, Fowler DW, Martin T, Pfretzschner H-U, & Ge S (2015) Dinosaur teeth from the Jurassic Qigu and Shishugou Formations of the Junggar Basin (Xinjiang/China) and their paleoecologic implications. Paläontologische Zeitschrift 89(3), p. 485-502. (Free Access PDF)

Scannella JB, Fowler DW, Goodwin MB, Horner JR (2014) Evolutionary trends in Triceratops from the Hell Creek Formation, Montana. Proceedings of the National Academy of Sciences 111(28), p. 10245-10250. (Free Access PDF)

Scannella JB, & Fowler DW (2014). A stratigraphic survey of Triceratops from the Hell Creek Formation, northeastern Montana (2006-2010). in Wilson et al. (eds) Through the End of the Cretaceous in the Type Locality of the Hell Creek Formation in Montana and Adjacent Areas: Geological Society of America Special Paper 503: p. 313-332. (Abstract & PDF request page)

Newbrey MG, Brinkman DB, Winkler DA, Freedman EA, Neuman AG, Fowler DW, & Woodward HN (2013) Teleost centrum and jaw elements from the Upper Cretaceous Nemegt Formation (Campanian – Maastrichtian) of Mongolia and a re-identification of the fish centrum found with the theropod Raptorex kreigsteini. pp. 291-303 in Arratia, Schultze & Wilson (eds), Mesozoic Fishes 5 - Global Diversity and Evolution. Verlag, Munchen, Germany. ISBN 978-3-89937-159-8. (Free Access PDF)

Woodruff DC, & Fowler DW (2012) Ontogenetic influence on neural spine bifurcation in Diplodocoidea (Dinosauria: Sauropoda): a critical phylogenetic character. Journal of Morphology 273(7), p. 754-764. (Abstract & PDF Request)

Koenig AE, Lucas SG, Neymark LA, Heckert AB, Sullivan RM, Jasinski SE, & Fowler DW (2012) Direct U-Pb dating of Cretaceous and Paleocene dinosaur bones, San Juan Basin, New Mexico. COMMENT, Geology 40: e262, doi:10.1130/G32154C.1 (Free Access Link)

Fowler DW, Freedman EA, Scannella JB, & Kambic RE (2011) The predatory ecology of Deinonychus and the origin of flapping in birds. PLoS One 6(12): e28964. doi:10.1371/journal.pone.0028964 (Free Access)

Fowler DW, & Sullivan RM (2011) The first giant titanosaurian sauropod from the Upper Cretaceous of North America. Acta Palaeontologica Polonica 56(4): 685-690. http://www.app.pan.pl/article/item/app20100105.html (Free Access)

Fowler DW, Woodward HN, Freedman EA, Larson PL, & Horner JR (2011) Reanalysis of “Raptorex kriegsteini”: a juvenile tyrannosaurid dinosaur from Mongolia. PLoS One 6(6): e21376. doi:10.1371/journal.pone.0021376 (Free Access).

Fowler DW, & Hall LE (2011) Scratch-digging sauropods, revisited, Historical Biology 23(1): 27-40. (Abstract and PDF Request)

Wings O, Schwarz-Wings D, & Fowler DW (2011). New sauropod material from the Late Jurassic part of the Shishugou Formation (Junggar Basin, Xinjiang, NW China). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 262/2: 129-150. (Free Access)

Sullivan RM, & Fowler DW (2011) Navajodactylus boerei, n.gen., n.sp. (Pterosauria, ?Azhdarchidae) from the Upper Cretaceous Kirtland Formation (Upper Campanian) of New Mexico. New Mexico Museum of Natural History and Science Bulletin 53: p. 393-404. (Free Access)

Fowler DW, Freedman EA, & Scannella JB (2009) Predatory functional morphology in raptors: Interdigital variation in talon size is related to prey restraint and immobilisation technique. PLoS One 4(11): e7999. doi:10.1371/journal.pone.0007999 (Free Access)

Lucas SG, Sullivan RM, Cather SM, Jasinski SE, Fowler DW, Heckert AB, Spielmann JA & Hunt AP (2009) No definitive evidence of Paleocene dinosaurs in the San Juan Basin. Paleontologica Electronica 12(2); 8A: 10p. (Free Access)

Wolff EDS, Fowler DW, & Bonde JW (2007) A possible case of necrotizing dermatitis in the crocodylian Diplocynodon, from the Oligocene of the Isle of Wight, UK. Historical Biology 19(2): p. 203-207. (Abstract and PDF Request)

Fowler DW, & Sullivan RM (2006) A ceratopsid pelvis with toothmarks from the Upper Cretaceous Kirtland Formation, New Mexico: Evidence of late Campanian tyrannosaurid feeding behavior. New Mexico Museum of Natural History and Science Bulletin 35: p. 127-130. (Free Access)

Sullivan RM, & Fowler DW (2006). New specimens of the rare ankylosaurid dinosaur Nodocephalosaurus kirtlandensis (Ornithischia: Ankylosauridae) from the Upper Cretaceous Kirtland Formation (De-na-zin Member), San Juan Basin, New Mexico. New Mexico Museum of Natural History and Science Bulletin 35: p. 259-261. (Free Access) 


Articles in edited books

Fowler DW (2017) Dinosaur evolution: it's about time. p40-41 in Cullum & Martinus (eds), 52 more things you should know about palaeontology. Agile Libre, Nova Scotia, Canada.


Conference Presentations & Extended Abstracts

Fowler DW, Wilson JP, Freedman Fowler EA, & Horner JR (2019) The Horned Dinosaur Leptoceratops (Ornithischia: Neoceratopsia) Raised its Young in Communal Nesting Burrows: Evidence from Three New Bonebeds in the Hell Creek Formation (Maastrichtian, Late Cretaceous), Montana. Abstracts volume of Cretaceous and Beyond, Paleontology of the Western Interior, Publications of North Dakota Geological Survey, Miscellaneous Series (94): 20.

Freedman Fowler EA & Fowler DW (2019) “Warwick's Duck", an Exceptional Specimen of Edmontosaurus (Dinosauria: Hadrosauridae) with Extensive Caudal Neural Spine Pathologies: Evidence of Mating Injuries in Hadrosaurs? Abstracts volume of Cretaceous and Beyond, Paleontology of the Western Interior, Publications of North Dakota Geological Survey, Miscellaneous Series (94): 21.

Canoy Illies, MM & Fowler DW (2019) Triceratops with a kink: Co-ossification of five distal caudal vertebraefrom the Hell Creek Formation of North Dakota.  Abstracts volume of Cretaceous and Beyond, Paleontology of the Western Interior, Publications of North Dakota Geological Survey, Miscellaneous Series (94): 11.

Fowler DW, Freedman Fowler EA & Alexander, JM (2018) The finest fossil owl. Journal of Vertebrate Paleontology 37(5, abstracts vol): 129.

Fowler DW & Freedman Fowler EA (2017) The oldest "Chasmosaurus"? A new skull from the Judith River Formation of Montana. Journal of Vertebrate Paleontology 36(5, abstracts vol): 114.

Fowler DW & Horner JR (2016) The evolutionary process of reinforcement reconciles conflicting functional hypotheses for bizarre structures in North American Late Cretaceous dinosaurs, explains apparent endemism as an artifact of vicariance and anagenesis, and suggests pre K-Pg diversity patterns may be peculiar to North America. Journal of Vertebrate Paleontology 35(5, abstracts vol): 138.

Woodruff DC & Fowler D (2016) What factors influence our reconstructions of Morrison Formation sauropod diversity? Journal of Vertebrate Paleontology 35(5, abstracts vol): 253.

Flora HM, Wilson JP, Gardner JD, & Fowler DW (2015) A three-dimensionally articulated probable oviraptorosaur from the Hell Creek Formation of Montana. Journal of Vertebrate Paleontology 34(5, abstracts vol): 124.

Anduza D & Fowler DW (2014) How might spinosaurids have caught fish? testing behavioral inferences through comparisons with modern fish-eating tetrapods. Journal of Vertebrate Paleontology, also presented at Mid Mesozoic, the age of dinosaurs in transition, Fruita CO.

Woodruff DC & Fowler DW (2014) The effects of ontogeny in regards to Morrison sauropod diversity. Mid Mesozoic, the age of dinosaurs in transition, Fruita CO: p. 81.

Fowler DW, Freedman-Fowler EA, Scannella JB, & Horner JR (2013) The influence of multi-niche ontogeny on differential survivorship across the K-Pg boundary. Journal of Vertebrate Paleontology 32(5, abstracts vol): 129.

Anduza D, Fowler DW, Noto C, & Horner JR (2013) New alvarezsaurid material from the Hell Creek Formation, Montana. Journal of Vertebrate Paleontology 32(5, abstracts vol): 77-78.

Woodruff C, Fowler DW, & Horner JR (2013) Changes in vertebral morphology associated with histologic data support significant change through ontogeny in diplodocid sauropods, Journal of Vertebrate Paleontology 32(5): 239.

Scannella JB, Fowler DW, Goodwin MG, & Horner JR (2013) The clandestine role of heterochrony in ceratopsian evolution as revealed by juvenile TriceratopsJournal of Vertebrate Paleontology 32(5): 206.

Fowler DW, Scannella JB, Goodwin MG, & Horner JR (2012) How to eat a Triceratops: large sample of toothmarks provides new insight into the feeding behavior of TyrannosaurusJournal of Vertebrate Paleontology 32(5): 96.

Scannella JB, Fowler DW, Goodwin MG, & Horner JR (2012) Transitional Triceratops: Details of an ontogenetic sequence from the upper Middle Unit of the Hell Creek Formation, Montana. Journal of Vertebrate Paleontology 32(5): 166.

Fowler DW (invited speaker), Freedman EA, Scannella JB, & Kambic RE (2011) The predatory ecology of Deinonychosauria: foot use compared among dromaeosaurids, troodontids and basal birds. Dinosaur Track Symposium, Obenkirchen, Germany: p. 22.

Fowler DW, Scannella JB, & Horner JR (2011) Reassessing ceratopsid diversity using unified frames of reference. Journal of Vertebrate Paleontology 31(5): 111.

Scannella JB, Fowler DW, Trevethan IJ, Roberts D, & Horner JR (2011) Individual variation in Triceratops from the Hell Creek Formation, Montana:, implications for dinosaur taxonomy. Journal of Vertebrate Paleontology 31(5): 187.

Sander PM, Wings O, Griebeler E, Fowler DW, & Henderson D (2011) High maximum growth rate in c.f. Mamenchisaurus argues against low basal metabolic rate as the explanation for sauropod gigantism. Journal of Vertebrate Paleontology 31(5): 186.

Fowler DW (2010) Anagenesis and long-term morphologic trends in Chasmosaurinae (Dinosauria: Ceratopsidae) revealed by a new high-resolution chronostratigraphic framework, ontogenetic analysis, and description of two new taxa. Journal of Vertebrate Paleontology 30(5): 91A.

Freedman EA, & Fowler DW (2010) Stratigraphic correlation of Judith River Formation (Campanian, Upper Cretaceous) exposures in Kennedy Coulee (Northcentral Montana) to the Foremost Formation (Alberta): Implications for anagenesis in hadrosaurid dinosaurs. Journal of Vertebrate Paleontology 30(5): 92A.

Padian K, Horner JR, Fowler DW, & Scannella JB (2010) How a synergy of species recognition and social signaling explains cranial anatomy and ontogeny in several groups of dinosaurs. Journal of Vertebrate Paleontology 30(5): 143A.

Woodruff DC, & Fowler DW (2010) Ontogenetic influence on neural spine bifurcation in Diplodocoidea (Dinosauria: Sauropoda): a critical phylogenetic character. Journal of Vertebrate Paleontology 30(5): 189A.

Fowler DW (2009) The grasping foot of Deinonychus: implications for predator ecology, evolution of the perching foot, and a new hypothesis for the origin of flight in birds. Journal of Vertebrate Paleontology 29(3): 98A.

Fowler DW (2009) A sequence stratigraphic subdivision of the Hell Creek Formation: beginnings of a high-resolution regional chronostratigraphic framework for the terminal Cretaceous. North American Paleontological Convention: p. 136.

Scannella JB, & Fowler DW (2009) Anagenesis in Triceratops: evidence from a newly resolved stratigraphic framework for the Hell Creek Formation. North American Paleontological Convention: p. 148.

Fowler DW, & Hall LE (2009) Scratch-digging sauropods, revisited: similarity of sauropod pes unguals to tortoises, and inferred nest-excavating behaviour. Museum of the Rockies occasional papers: Dinosaur Eggs & Babies: p. 18.

Wings O, Fowler DW, Maisch MW, Martin T, Pfretzschner H-U & Sun, G (2007) Dinosaur teeth from the Jurassic Qigu and Shishugou Formations of the Junggar Basin, Xinjiang/China and their palaeobiogeographical implications. In: Sun, G; Mosbrugger, V.; Sun, Y.W. & Bruch, A. (eds.) - Proceedings of the International Symposium for Sino-German Cooperation on Geology and Environmental Changes in Northern China; Urumqi/China: p. 5-10.

Fowler DW (2007) Recently rediscovered baryonychine teeth (Dinosauria: Theropoda): new morphologic data, range extension, & similarity to CeratosaurusJournal of Vertebrate Palaeontology 27(3): 76A.

Wings O, Sander PM, Tutken T, Fowler DW, & Sun G (2007) Growth and life history of Asia's largest dinosaur. Journal of Vertebrate Palaeontology 27(3): 167A.

Fowler DW (2006) Terrestrial Late Cretaceous stratigraphy of North America and the utility of ceratopsids in biostratigraphy. Journal of Vertebrate Palaeontology 26(3): 63A. (poster)

Fowler DW, Simmonds K, Green M, & Stevens KA (2003) The taphonomic setting of two mired sauropods (Wessex Fm, Isle of Wight, UK), palaeoecological implications and taxon preservation bias in a Lower Cretaceous wetland. Journal of Vertebrate Palaeontology 23(3): 51A. (poster)


Book Reviews

Fowler DW (2017) Species and speciation in the fossil record. Edited by Allmon & Yacobucci. Chicago (Illinois): University of Chicago Press. Reviewed in: The Quarterly Review of Biology 92(4), p. 461.

Fowler DW (2013) Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems. Life of the Past. Edited by Pascal Godefroit. Bloomington (Indiana): Indiana University Press. Reviewed in: The Quarterly Review of Biology 88(3), p. 235.